1. Department of Biomedical Engineering Pennsylvania State University University Park PA 16802‐4400 USA
2. Department of Veterinary and Biomedical Sciences Pennsylvania State University University Park PA 16802‐4400 USA
3. Center for Infectious Disease Dynamics Pennsylvania State University University Park PA 16802‐4400 USA
4. Center for Molecular Immunology and Infectious Disease Pennsylvania State University University Park PA 16802‐4400 USA
5. Huck Institutes of the Life Sciences Penn State University University Park PA 16802‐4400 USA