1. Center for Zoo and Wild Animal Health; Copenhagen Zoo; Roskildevej 32 2000 Frederiksberg Denmark
2. Faculty of Health and Medical Sciences; Department of Veterinary Disease Biology; University of Copenhagen; Stigboejlen 4 1870 Frederiksberg C Denmark
3. Faculty of Science and Technology; Department of Bioscience; Arctic Research Centre (ARC); Aarhus University; Frederiksborgvej 399 PO Box 358 4000 Roskilde Denmark