1. University of Cambridge, Department of Physiology, Development & Neuroscience; Downing Street; Cambridge CB2 3EG United Kingdom
2. Department of Zoology and Entomology; University of Pretoria; Pretoria 0002 South Africa
3. Sorbonne Universités, CR2P, UMR 7207 du CNRS, Département Histoire de la Terre, Muséum National d'Histoire Naturelle et Université Pierre et Marie Curie; France