1. Department of Botany and Zoology Stellenbosch University Private Bag X1 Matieland 7602 South Africa
2. H. G. W. J. Schweickerdt Herbarium, Department of Plant and Soil Sciences, Plant Sciences Complex University of Pretoria Private Bag X20 Hatfield 0028 South Africa
3. Institute of Botany, Academy of Sciences of the Czech Republic, Průhonice, Czech Republic, and Department of Botany Charles University Praha Czech Republic