1. Department of Biology, Faculty of Forestry Canadian Rivers Institute University of New Brunswick Post Office Box 4400 Fredericton New Brunswick E3B 5A3 Canada
2. Department of Forestry Canadian Rivers Institute University of New Brunswick Post Office Box 4400 Fredericton New Brunswick E3B 5A3 Canada
3. Department of Biology School of Earth Environment and Society McMaster University 1280 Main Street West Hamilton Ontario L8S 4K1 Canada