1. Department of Biology University of Hawai'i at Hilo 200 West Kawili Street Hilo HI96712USA
2. National Park Service, Inventory and Monitoring Program, Pacific Island Network P.O. Box 52 Hawai'i Volcanoes National Park HI96718USA
3. Department of Geography and Environmental Studies University of Hawai'i at Hilo 200 West Kawili Street Hilo HI96712USA
4. Department of Biological Sciences National University of Singapore 14 Science Drive 4117543Singapore