1. Institute of Low Temperature Science, Hokkaido University; Japan
2. Graduate School of Environmental Science; Hokkaido University; Japan
3. Antarctic Climate and Ecosystems Cooperative Research Centre, University of Tasmania; Hobart Tasmania Australia
4. National Institute of Polar Research; Tokyo Japan
5. Now at Marine Works Japan; Yokohama Japan
6. CSIRO Oceans and Atmosphere; Hobart Tasmania Australia
7. SOKENDAI, Graduate University for Advanced Studies; Tokyo Japan