1. Key Laboratory for Experimental Teratology of Ministry of Education and Department of Immunology; Shandong University School of Basic Medical Science; Jinan Shandong, 250012 People's Republic of China
2. Key Laboratory of Infection and Immunity of Shandong Province, School of Basic Medical Sciences; Shandong University; Jinan People's Republic of China
3. Key Laboratory for Experimental Teratology of Ministry of Education and Department of Genetics; Shandong University School of Basic Medical Science; Jinan Shandong, 250012 People's Republic of China
4. Department of Microbiology, Immunology and Molecular Genetics; Lexington KY
5. Markey Cancer Center, University of Kentucky College of Medicine; Lexington KY