1. Jigme Dorji National Park Department of Forests and Park Services Ministry of Agriculture and Forests Royal Government of Bhutan Gasa Bhutan
2. Durrell Institute of Conservation and Ecology School of Anthropology and Conservation University of Kent Canterbury UK
3. Wildlife Science Unit School of Agriculture and Food Sciences The University of Queensland Gatton Qld Australia
4. Nature Conservation Division Department of Forests and Park Services Ministry of Agriculture and Forests Royal Government of Bhutan Thimphu Bhutan
5. Wildlife Conservation Research Unit Department of Zoology The Racanati‐Kaplan Centre University of Oxford Tubney UK