1. Smithsonian Environmental Research Center, Fisheries Conservation Laboratory 647 Contees Wharf Road Edgewater Maryland21037USA
2. National Marine Fisheries Service, Highly Migratory Species Management Division Gloucester Massachusetts01930USA
3. Chesapeake Biological Laboratory University of Maryland Center for Environmental Science Solomons Maryland20688USA
4. Virginia Institute of Marine Science, College of William and Mary Gloucester Point Virginia23062USA