1. Department of Animal and Plant Sciences; University of Sheffield; Sheffield S10 2TN United Kingdom
2. Department of Archaeology, Northgate House; University of Sheffield; Sheffield S1 4ET United Kingdom
3. Laboratoire Évolution and Diversité Biologique (EDB UMR5174); Université de Toulouse; CNRS, ENSFEA, IRD, UPS France
4. Current Address: School of Archaeology; University of Oxford; 36 Beaumont Street Oxford OX1 2PG United Kingdom