1. School of Anatomical Sciences, Faculty of Health Sciences University of the Witwatersrand Johannesburg South Africa
2. Division of Clinical Anatomy and Biological Anthropology, Department of Human Biology University of Cape Town Cape Town South Africa
3. Centre for Zoo and Wild Animal Health Copenhagen Zoo Frederiksberg Denmark
4. Borås Zoo Borås Sweden
5. Department of Anthropology and Center for the Advanced Study of Human Paleobiology The George Washington University Washington District of Columbia USA