1. Department of Wildlife, Fisheries, and Aquaculture, Mississippi State University MS USA
2. Quantitative Ecology and Spatial Technologies Laboratory, Mississippi State University MS USA
3. Geosystems Research Institute, Mississippi State University MS USA
4. School of Mathematics and Statistics, University of Sheffield Sheffield UK
5. Department of Biosciences, Swansea University Swansea UK
6. Centre for Biomathematics, Swansea University Swansea UK