1. Behavior; Ecology; Evolution and Systematics Section; School of Biological Sciences; Illinois State University; Normal; 61790-4120; Illinois
2. Department of Biological Sciences; Grant MacEwan University; Edmonton; Alberta; T5J 4S2; Canada
3. Centre for Ecology & Conservation; School of Biosciences; University of Exeter in Cornwall; Cornwall; Penryn; TR10 9EZ; U.K