1. Sorbonne Universités, UPMC University Paris 06, CNRS, UMR 8227; Integrative Biology of Marine Models, Station Biologique Roscoff; CS 90074, 29688 Roscoff Brittany France
2. University of Brest, CNRS, IRD; Ifremer, LEMAR UMR 6539, IUEM, Place Nicolas Copernic; 29280 Plouzané Brittany France