1. Key Laboratory of Aquaculture in South China Sea for Aquatic Economic Animal of Guangdong Higher Education Institutes College of Fishery, Guangdong Ocean University Zhanjiang China
2. Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Economic Animals College of Fishery, Guangdong Ocean University Zhanjiang China
3. Faculty of Chemistry and Environmental Science Guangdong Ocean University Zhanjiang China
4. State Laboratory of Developmental Biology of Freshwater Fish College of Life Sciences, Hunan Normal University School Changsha China