1. Graduate School of Fisheries and Environmental Sciences Nagasaki University, Nagasaki‐shi Nagasaki Japan
2. Atmospheric and Ocean Research Institute the University of Tokyo, Kashiwa‐shi Chiba Japan
3. Tokyo University of Marine Science and Technology, Minato‐ku Tokyo Japan
4. Yokohama Field Station, Fisheries Resources Institute Japan Fisheries Research and Education Agency, Yokohama‐shi Kanagawa Japan
5. Graduate School of Science and Engineering Soka University, Hachioji‐shi Tokyo Japan