1. Department of Ecology and Evolutionary Biology University of California Los Angeles California 90095 USA
2. Forestry and Forest Products Research Institute Tsukuba Ibaraki 305‐8687 Japan
3. Yale School of Forestry and Environmental Studies Yale University New Haven Connecticut 06511 USA
4. Smithsonian Tropical Research Institute Unit 0948 Panama City Panama
5. Field Museum of Natural History Chicago Illinois 60605 USA
6. Morton Arboretum Lisle Illinois 60532 USA