1. Program in Ecology, Evolution and Conservation Biology Department of Natural Resources and Environmental Science University of Nevada Reno Mail Stop 186 Reno Nevada 89557 USA
2. Department of Fish and Wildlife Conservation Virginia Polytechnic Institute and State University Blacksburg Virginia 24060 USA
3. Department of Wildlife, Fisheries, and Conservation Biology University of Maine 5755 Nutting Hall Room 210 Orono Maine 04469 USA