1. Graduate School of Pharmaceutical Sciences The University of Tokyo 7-3-1 Hongo Bunkyo-ku Tokyo 113-0033 Japan
2. Collaborative Research Institute for Innovative Microbiology The University of Tokyo Yayoi 1-1-1, Bunkyo-ku Tokyo 113-8657 Japan
3. PRESTO (Japan) Science and Technology Agency Kawaguchi Saitama 332-0012 Japan