1. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor Michigan 48109 USA
2. Department of Biology University of Maryland College Park Maryland 20742 USA
3. Department of Biology University of Puerto Rico Río Piedras Puerto Rico 00931 USA
4. Centre for Ecology & Hydrology Bush Estate, Penicuik Midlothian EH26 0QB United Kingdom
5. Department of Ecology, Evolution & Environmental Biology Columbia University New York New York 10027 USA
6. Department of Environmental Sciences University of Puerto Rico Río Piedras Puerto Rico 00936 USA
7. Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Kunming China