1. Department of Microbiology and Infection Faculty of Medicine Kochi University Nankoku City Kochi Japan
2. Center for Innovative and Translational Medicine Faculty of Medicine Kochi University Nankoku City Kochi Japan
3. Research Institute of Molecular Genetics Kochi University Nankoku City Kochi Japan
4. Science Research Center Kochi University Nankoku City Kochi Japan
5. Research Center for Infections and Antimicrobials Kitasato Institute for Life Sciences Kitasato University Tokyo Japan