1. School of Grassland Science, Beijing Forestry University, 35 Qinghua East Road, Beijing 100083, China
2. Department of Plant Biology, Rutgers, the State University of New Jersey, 59 Dudley Road, New Brunswick, NJ 08901, USA
3. Department of Plant Biology, Rutgers, the State University of New Jersey, Rutgers Agricultural Research and Extension Center, Bridgeton, NJ 08302, USA
4. Department of Horticulture, Oregon State University, 4017 Agriculture and Life Sciences Building, Corvallis, OR 97331, USA