1. College of Horticulture, Hainan University, Haikou 570228, Hainan, China
2. Tropical Crops Genetic Resources Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou 571737, Hainan, China
3. Multidisciplinary Workshop on Vascular Plants, Border Ecology Laboratory, University of Flores, Sede Comahue (UFLO), Rio Negro, Argentina