1. Department of Zoology, University of Sri Jayewardenepura, Gangodawila, Nugegoda, Sri Lanka
2. Inland Aquatic Resources and Aquaculture Division, National Aquatic Resources Research and Development Agency (NARA), Crow Island, Colombo 15, Sri Lanka
3. Department of Food Science and Technology, University of Sri Jayewardenepura, Gangodawila, Nugegoda, Sri Lanka