1. Department of Microbiology, School of Medicine, Iran University of Medical Sciences, Tehran, Iran
2. School of Medicine, Shahid Beheshti University of Medical Sciences, Tehran, Iran
3. Razi Vaccine and Serum Research Institute, Agricultural Research, Education and Extension Organization (AREEO), Karaj, Iran
4. Department of Microbiology, School of Medicine, Shahid Beheshti University of Medical Sciences, Tehran, Iran