1. a School of Agriculture, Policy and Development, University of Reading , Whiteknights, PO Box 237, Reading, UK
2. b Plant Production and Protection Division , Food and Agriculture Organization (FAO) of the United Nations , Viale delle Terme di Caracalla, 00153, Rome, Italy
3. c Tropical Agriculture Association , PO Box 3, Penicuik, Midlothian, EH26 0RX, UK
4. d Department of Biological Sciences , University of Essex , Wivenhoe Park, Colchester, Essex, CO4 3SQ, UK