1. School of Biosciences, University of Birmingham, Birmingham B15 2TT, United Kingdom
2. Division of Plant and Crop Sciences, School of Biosciences and Centre for Plant Integrative Biology, University of Nottingham, Loughborough LE12 5RD, United Kingdom
3. Department of Plant Biology and Genome Center, University of California, Davis, California 95616
4. Department of Plant Biology and Forest Genetics, Uppsala BioCenter, Swedish University of Agricultural Sciences and Linnean Center for Plant Biology, SE-75007 Uppsala, Sweden
5. Department of Plant Biology, Carnegie Institution for Science, Stanford, California 94305