1. Microbial and Environmental Genomics, J. Craig Venter Institute, La Jolla, California 92037
2. Integrative Oceanography Division, Scripps Institution of Oceanography, UC San Diego, La Jolla, California 92037
3. Department of Biology, Washington University, St. Louis, Missouri 63130
4. Institut de Biologie de l’École Normale Supérieure, École Normale Supérieure, Paris Sciences et Lettres Research University, CNRS UMR8197 INSERM U1024, 75005 Paris, France