1. National Institute of Agrobiological Resources, Tsukuba, Ibaraki 305-8602, Japan
2. Bio-oriented Technology Research Advancement Institution, Omiya, Saitama 331-8537, Japan
3. Institute of the Society for Techno-innovation of Agriculture, Forestry and Fisheries, Tsukuba, Ibaraki 305-0854, Japan
4. Department of Biology, Faculty of Science, Toyama University, Toyama, 930-8555, Japan