1. Integrative Cell Biology Laboratory, School of Biological and Biomedical Sciences, University of Durham, South Road, Durham DH1 3LE, United Kingdom
2. Institute of Applied Genetics and Cell Biology, BOKU, University of Natural Resources and Applied Life Sciences, Muthgasse 18 A-1190 Vienna, Austria
3. Himeji Institute of Technology, Faculty of Science, Hyogo, Japan
4. Department of Cell and Developmental Biology, John Innes Centre, Norwich NR4 7UH, United Kingdom