1. Department of Biochemistry and Molecular Biology, Michigan State University, East Lansing, Michigan 48824
2. Department of Energy–Plant Research Laboratory, Michigan State University, East Lansing, Michigan 48824
3. Cell and Molecular Biology Program, Michigan State University, East Lansing, Michigan 48824
4. Genetics Program, Michigan State University, East Lansing, Michigan 48824
5. Department of Botany, University of Toronto, Toronto, Ontario, Canada M5S 3B2