1. Department of Biology and the Huck Institutes of the Life Sciences, Pennsylvania State University, University Park, Pennsylvania 16802
2. Intercollege Graduate Program in Cell and Developmental Biology, Pennsylvania State University, University Park, Pennsylvania 16802
3. College of Biological Science and Biotechnology, Beijing Forestry University, Beijing 100083, China
4. State Key Laboratory of Genetic Engineering and Institute of Plant Biology, Institute of Genetics, Center for Evolutionary Biology, School of Life Sciences, Fudan University, Shanghai 200433, China