1. Department of Cell Biology, National Institute for Basic Biology, Okazaki 444-8585, Japan
2. Department of Basic Biology, School of Life Science, The Graduate University for Advanced Studies, Okazaki 444-8585, Japan
3. Institut National de la Recherche Agronomique, UMR1318, Institut Jean-Pierre Bourgin, RD10, F-78000 Versailles, France
4. AgroParisTech, Institut Jean-Pierre Bourgin, RD10, F-78000 Versailles, France
5. Institute of Plant Science and Resources, Okayama University, Okayama 710-0046, Japan
6. Frontier Research Center, Tokyo Institute of Technology, Yokohama 226-8503, Japan