1. Department of Biotechnology and Life Science, Faculty of Engineering, Tokyo University of Agriculture and Technology, Koganei, Tokyo 184-8588, Japan
2. Department of Agricultural Research, Ehime Research Institute of Agriculture, Forestry and Fisheries, Matsuyama, Ehime 799-2405, Japan
3. Faculty of Environmental Horticulture, Minami Kyushu University, Miyakonojo, Miyazaki 885-0035, Japan