1. Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China
2. Key Laboratory of Plant Stress Biology, School of Life Sciences, Henan University, Kaifeng 457004, China
3. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design and College of Biological Sciences and Biotechnology, Beijing Forestry University, Beijing 100083, China
4. State Key Laboratory of Plant Cell and Chromosome Engineering, Centre for Molecular Agrobiology, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing100101, China
5. Institute of Cellular and Molecular Botany, Rheinische Friedrich-Wilhelms-University Bonn, Department of Plant Cell Biology, Bonn D-53115, Germany
6. Centre of the Region Hana for Biotechnological and Agricultural Research, Faculty of Science, Palacky University, Olomouc 78301, Czech Republic