1. Sainsbury Laboratory, John Innes Centre, Norwich NR4 7UH, United Kingdom
2. Commissariat à l'Energie Atomique (CEA), Institut de Biologie et Technologies de Saclay (IBITECS), and Centre National de la Recherche Scientifique (CNRS), Gif-sur-Yvette F-91191, France
3. RIKEN Plant Science Center, Tsurumi-ku, Yokohama, 230-0045 Japan
4. School of Chemical Sciences and Pharmacy, University of East Anglia, Norwich NR4 7TJ, United Kingdom
5. Department of Biology, University of York, York YO10 5YW, United Kingdom