1. Graduate School of Science and Technology, Shizuoka University, Shizuoka 422-8529, Japan
2. Faculty of Agriculture, Shizuoka University, Shizuoka 422-8529, Japan
3. Department of Biology, Lakehead University, Thunder Bay, Ontario, P7B 5E1 Canada
4. Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany
5. Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin 541006, China