1. Laboratory for Morphogenetic Signaling, RIKEN Center for Developmental Biology, Kobe, Hyogo 650-0047, Japan
2. Department of Biology, Kobe University Graduate School of Science, Nada-ku, Hyogo 657-8501, Japan
3. Graduate School of Biostudies, Kyoto University, Sakyo-ku 606-8501, Japan
4. The Keihanshin Consortium for Fostering the Next Generation of Global Leaders in Research (K-CONNEX), Sakyo-ku, Kyoto 606-8501, Japan