1. The Fenner School of Environment and Society; The Australian National University; Canberra Australian Capital Territory 2601 Australia
2. School of Life and Environmental Sciences; Centre for Integrative Ecology; Deakin University Geelong; Melbourne Burwood Campus Melbourne Victoria 3125 Australia
3. Long-term Ecological Research Network; Terrestrial Ecosystems Research Network; The Australian National University; Canberra Australian Capital Territory 2601 Australia