1. Department of Ecology and Natural Resource Management; Norwegian University of Life Sciences; 1430 Ås Norway
2. Faculty of Biological Sciences; University of Leeds; Leeds LS2 9JT UK
3. Instituto de Investigación de Recursos Biológicos Alexander von Humboldt; Calle 28A # 15-09 Bogotá Colombia
4. Department of Animal and Plant Sciences; University of Sheffield; Sheffield S10 2TN UK
5. School of Marine & Tropical Biology; James Cook University; Cairns Queensland 4811 Australia