1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan 650201 China
2. Institute of Tibetan Plateau Research at Kunming; Chinese Academy of Sciences; Kunming Yunnan 650201 China
3. Plant Germplasm and Genomics Center; Germplasm Bank of Wild Species in Southwest China; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan 650201 China
4. Key Laboratory of Plant Resources Conservation and Utilization and College of Biology and Environmental Sciences; Jishou University; Jishou Hunan 416000 China
5. Department of Botany; National Museum of Natural History; MRC 166; Smithsonian Institution; Washington DC 20013-7012 USA