1. State Key Laboratory of Biocatalysis and Enzyme Engineering, School of Life Sciences Hubei University Wuhan 430062 China
2. National Key Laboratory for Germplasm Innovation & Utilization of Horticultural Crops, College of Horticulture and Forestry Sciences Huazhong Agricultural University Wuhan 430070 China
3. Hubei Hongshan Laboratory Wuhan 430070 China
4. School of Plant Sciences, College of Agriculture and Life Sciences The University of Arizona Tucson Arizona 85721 USA
5. College of Life Sciences Shandong Normal University Jinan 250014 China