1. United Graduate School of Agricultural Sciences; Kagoshima University; Kagoshima Japan
2. Faculty of Fisheries; Kagoshima University; Kagoshima Japan
3. Research Fellow of Japan Society for the Promotion of Science; Tokyo Japan
4. Saga Prefectural Ariake Fisheries Research and Development Center (SAFREDEC); Saga Japan
5. Institute for East China Sea Research, Organization for Marine Science and Technology; Nagasaki University; Nagasaki Japan