1. Department of Biological Sciences; Dornsife College of Letters, Arts and Sciences; University of Southern California; Los Angeles California 90089 USA
2. Biosciences; College of Life and Environmental Sciences; University of Exeter; Exeter EX4 4QD UK
3. Département de Biologie; Takuvik Joint International Laboratory; Centre National de la Recherche Scientifique (France, CNRS UMI 3376); Québec Océan; Institut de Biologie Intégrative et des Systèmes (IBIS); Université Laval; Québec Quebec Canada