1. Department of Entomology; Center for Pollinator Research; The Pennsylvania State University; University Park PA 16802 USA
2. School of Biological Sciences; Royal Holloway University of London; Egham TW 20 0EX UK
3. Institut de Recherche sur la Biologie de l‘Insecte (UMR 7261), CNRS; University of Tours; Parc de Grandmont 37200 Tours France
4. Department of Ecology & Evolution; University of Lausanne; Biophore Unil-Sorge Lausanne Switzerland
5. USDA-ARS; Gainesville FL 32608 USA