1. Department of Ecology and Evolutionary Biology University of Connecticut Storrs Connecticut USA
2. Universidad Nacional Laboratorio de Ecología Funcional y Ecosistemas Tropicales, Escuela de Ciencias Biológicas Universidad Nacional Heredia Costa Rica
3. Department of Biology University of Miami Miami Florida USA
4. Fairchild Tropical Botanic Garden Coral Gables Florida USA
5. Department of Biological Sciences Humboldt State University Arcata California USA
6. Department of Botany, National Museum of Natural History, MRC‐166 Smithsonian Institution Washington District of Columbia USA
7. Escuela de Ciencias Agrarias Universidad Nacional Heredia Costa Rica