1. Department of Animal Ecology and Ecophysiology Institute for Water and Wetland Research Radboud University P.O. Box 9010 6500 GL Nijmegen the Netherlands
2. Marine Biology and Ecology Research Centre School of Marine Science and Engineering University of Plymouth Davy Building Drake Circus Plymouth PL4 8AA UK
3. Department of Evolution, Ecology & Behaviour Biosciences Building Institute of Integrative Biology University of Liverpool Liverpool L69 7ZB UK