1. School of Environmental Sciences University of Liverpool Liverpool UK
2. Department of Geography UCL London UK
3. Department of Zoology & Entomology University of Pretoria Pretoria South Africa
4. Soil Biodiversity Group, Department of Life Sciences Natural History Museum London UK
5. School of Biological and Behavioural Sciences Queen Mary University of London London UK
6. School of Animal, Plant and Environmental Sciences University of the Witwatersrand South Africa