1. State Key Laboratory of Marine Environmental Science; Xiamen University; Xiamen China
2. Institute of Marine Microbes and Ecospheres; Xiamen University; Xiamen China
3. Division of Life Science/Coastal Marine Laboratory; Hong Kong University of Science and Technology; Kowloon Hong Kong China
4. Microbial Ecology and Biogeochemistry Group; Laboratoire d'Océanographie de Villefranche; Université Pierre et Marie Curie-Paris 6; Villefranche-sur-Mer France
5. CNRS; Laboratoire d'Océanographie de Villefranche; Villefranche-sur-Mer France